In addition to estrogen, the granulosa cells also secrete a hormone called activin, which stimulates FSH production, as well as binding to FSH receptors, and the activity of granulosa cell aromatase as well.

除了雌激素，颗粒细胞还会分泌一种激素叫做激活素，它可以刺激FSH的分泌，与FSH受体结合；并且与颗粒细胞芳香酶的活性有关。