So the discovery of this creature kicked off a debate about which major taxonomic group, or clade, of reptiles that turtles belonged to.

因此，这种生物的发现引发了关于海龟属于哪个主要爬行动物分类群或进化枝的争论。

One theory holds that, if Rhyniognatha was a flying insect, that would mean that the pterygotes -- the taxonomic group that includes flying insects -- first came about in the early Devonian, say, more than 400 million years ago.

一种理论认为， 如果鼻颚龙是一种飞行昆虫，那就意味着翼龙——包括飞行昆虫在内的分类群——首先出现于泥盆纪早期，比方说，4 亿多年前。