Fretwell told the Associated Press that 30 percent of the 62 known penguin colonies were hurt by low levels of sea ice.

弗雷特威尔告诉美联社，在已知的62个企鹅栖息地中，有30%受到了海冰减少的影响。

The study noted that extremely low levels of sea ice in 2016 led to a large reproduction failure of an Emperor penguin group in Antarctica's Halley Bay.

该研究指出，2016年极低的海冰水平导致南极洲哈雷湾的帝企鹅种群大规模繁殖失败。