Indirect competition can be measured by the success of the alumni of each university.

间接竞争可以用每个大学校友的成功指数来衡量。

They can also harm other wild species through indirect competition rather than eating them or shrinking their food supply.

它们也可以通过间接竞争而不是吃掉它们或减少食物来伤害其他野生物种。

Though they were born from the same root, there have been many direct and indirect competitions between the two universities all the time.

虽然它们本是同根生，这两所大学之间一直都存在着直接或间接的竞争。