They carried with them diseases like measles, influenza, and smallpox that their people had gradually acquired genetic resistance to over thousands of years of dense civilization, agriculture, and livestock domestication.

他们携带了麻疹、流感和天花等疾病，他们的人民在数千年的密集文明、农业和牲畜驯化中逐渐获得了对这些疾病的遗传抵抗力。