Coordinate transformations are a basic calculation that animals need to perform to interact with the world.

坐标变换是动物与世界互动所需要进行的基本计算。

In the model, dragonflies do coordinate transformations in only one computational step, one layer of neurons.

在该模型中，蜻蜓只用一个计算步骤，即一个神经元层来做坐标转换。

If the responses that we record in the brain match those predicted by the model, we will have identified which neurons are responsible for coordinate transformations.

如果我们在大脑中记录的反应与模型预测的反应相匹配，我们就会确定哪些神经元负责坐标转换。

In other words, if I want to study how the dragonfly does coordinate transformations, the neural circuit that I need to understand, the neural circuit that I need to study, can have at most four layers of neurons.

换句话说，如果我想研究蜻蜓如何进行坐标变换，我需要了解神经回路，我需要研究神经回路，最多可以有四层神经元。