Also abundant were mayflies and caddis flies. These invertebrates are indicators of a healthy aquatic ecosystem.

也有大量的蜉蝣和石蛾。这些无脊椎动物是健康水生生态系统的标志。

Also abundant were mayflies and caddis flies.

蜉蝣和石蚕也很丰富。

This requires a more inclusive, and generous, definition of creativity, for it is not a trait obviously exhibited by a prairie dog or caddis fly.

这就需要对创造力有一个更包容广阔的定义，毕竟这可不是草原土拨鼠或石蛾能够明显表现出的特征。