He divided the mosses into six genera: Mnium, Hypnum, Polytrichum, Bryum, Sphagnum and Lycopodium — classifications that are useful today.

他把藓类分为六个属：提灯藓属、灰藓属、金发藓属、真藓属、泥炭藓属和石松属，这些分类在今天仍然有用。