But another cost may rise over time: the underdevelopment of human capital.

但随着时间的推移，另一个成本可能会上升：人力资本的发展不足。

The new state of South Sudan , which combines oil wealth with instability and underdevelopment, will surely join its ranks.

石油为苏丹南部的新政权带来财富，但那里依旧属于欠发达地区，并且不稳定。毫无疑问，它将会加入这一行列。

Delaying or inhibiting the arrival of these cells in certain areas of the body can result in an underdevelopment of the pituitary and adrenal glands, which govern fear and aggression.

延迟或抑制这些细胞到达身体的某些区域会导致脑垂体和肾上腺发育不全，这些腺体控制着恐惧和攻击性。

Therefore, Cape Town's water system relied on just a single critical point of failure that had been exacerbated by decades of population growth accompanied by underdevelopment of increasing the water supply.

因此，开普敦的供水系统仅仅依赖于一个关键的故障点，而这个故障点由于几十年来的人口增长和增加水供应的不发达而变得更加严重。