Discovery of subspecies toxic to dipteran and coleopteran insects in 1977 and 1983, respectively.

1977年以前只发现13个亚种，而且这些亚种都只对鳞翅目害虫有效。

Any of various insect larvae that live and feed within a leaf, including caterpillars, sawfly larvae, beetle and weevil grubs, and dipteran maggots.

许多在叶中生活并取食的昆虫幼虫的统称，包括毛虫、叶蜂幼虫、甲虫和象甲虫以及双翅类昆虫的蛆。