Fish began their transition onto land 400 million years ago, during in the Devonian period.

4 亿年前，在泥盆纪时期，鱼类开始向陆地过渡。

By the end of the Devonian period, plants had developed almost all of  the traits they would need to thrive.

到泥盆纪末期，植物已经发展出它们茁壮成长所需的几乎所有特征。